|
| ||||
Copyright © 2008 Ecostudies Institute
| A nonprofit organization committed to ecological research and conservation |
| ECOSTUDIES INSTITUTE |
| ECOSTUDIES INSTITUTE |
| Contact Information: Ecostudies Institute P.O. Box 703 Mount Vernon, WA 98273 305-213-8829 ecostudies@ecoinst.org |
Some files on this page
require Adobe Acrobat
Reader software. Click
on the image above to
download it for free
from the Adobe site.
require Adobe Acrobat
Reader software. Click
on the image above to
download it for free
from the Adobe site.
John Lloyd has been involved in research on birds since his days as an undergraduate at the University
of Vermont. John received his Ph.D. in wildlife biology from the University of Montana in 2003, where he
worked with Dr. Tom Martin on a study of the ecology, behavior, and conservation biology of Chestnut-
collared Longspurs (see more in Grassland Bird Conservation). Dr. Lloyd’s study of Chestnut-collared
Longspurs was highlighted in an article in Birding, the journal of the American Birding Association. More
recently, Dr. Lloyd has worked on finding scientific solutions to the problems that roads pose for
wildlife. Other current projects, highlighted on our project pages, include studies of the effects of fire on
birds in south Florida pine forests; taxonomy and population size of the Grand Bahama race of the
Brown-headed Nuthatch, an island endemic; and habitat requirements and population abundance of
birds in Florida mangrove forests. In all facets of his work, Dr. Lloyd is especially interested in using
ecological theory to understand and solve conservation problems.
of Vermont. John received his Ph.D. in wildlife biology from the University of Montana in 2003, where he
worked with Dr. Tom Martin on a study of the ecology, behavior, and conservation biology of Chestnut-
collared Longspurs (see more in Grassland Bird Conservation). Dr. Lloyd’s study of Chestnut-collared
Longspurs was highlighted in an article in Birding, the journal of the American Birding Association. More
recently, Dr. Lloyd has worked on finding scientific solutions to the problems that roads pose for
wildlife. Other current projects, highlighted on our project pages, include studies of the effects of fire on
birds in south Florida pine forests; taxonomy and population size of the Grand Bahama race of the
Brown-headed Nuthatch, an island endemic; and habitat requirements and population abundance of
birds in Florida mangrove forests. In all facets of his work, Dr. Lloyd is especially interested in using
ecological theory to understand and solve conservation problems.
Dr. Lloyd has published numerous articles in leading scientific journals such as the Journal of Wildlife
Management, Proceedings of the Royal Society of London, and Behavioral Ecology. John frequently
gives presentations at scientific meetings, and as an invited speaker has discussed topics ranging from
the statistical analysis of nest survival data to methods for reducing collisions between animals and
motor vehicles. In addition to his research collaborations with land managers and scientists in academia,
NGOs, and federal and state agencies, Dr. Lloyd has also served as a consultant to the Oregon
Department of Transportation and the USDA Forest Service, helping them develop strategies for making
highways less dangerous for wildlife.
Management, Proceedings of the Royal Society of London, and Behavioral Ecology. John frequently
gives presentations at scientific meetings, and as an invited speaker has discussed topics ranging from
the statistical analysis of nest survival data to methods for reducing collisions between animals and
motor vehicles. In addition to his research collaborations with land managers and scientists in academia,
NGOs, and federal and state agencies, Dr. Lloyd has also served as a consultant to the Oregon
Department of Transportation and the USDA Forest Service, helping them develop strategies for making
highways less dangerous for wildlife.
PUBLICATIONS
Some of Dr. Lloyd’s publications are available below in PDF format. For a complete list of publications and
presentations, see Dr. Lloyd’s CV.
Lloyd, J. D., and G. L. Slater. 2007. Environmental factors affecting productivity of brown-headed nuthatches.
Journal of Wildlife Management 71:1968-1975. PDF
Lloyd, J. D., and J. J. Tewksbury. 2007. Analyzing avian nest survival in forests and grasslands: a comparison of the
Mayfield and logistic-exposure methods. Studies in Avian Biology 34:96-104. PDF
Tewksbury, J. J., L. Garner, S. Garner, J. D. Lloyd, V. Saab, and T. E. Martin. 2006. Tests of landscape influence: nest
predation and brood parasitism in fragmented ecosystems. Ecology 87:759-76. PDF
Lloyd, J. D., and A. Casey. 2005. Wildlife hot spots along highways in northwestern Oregon. Final Report to the
Oregon Department of Transportation. PDF
Lloyd, J. D. and T.E. Martin. 2005. Reproductive success of Chestnut-collared Longspurs in native and exotic
grassland. Condor 107:363-374. PDF
Lloyd, J. D., and T. E. Martin. 2004. Nest-site preference and maternal effects on offspring growth. Behavioral
Ecology 15:816-823. PDF
Ruediger, B., and J. D. Lloyd. 2003. A rapid assessment protocol for identifying potential habitat linkages along
roads. Pages 205-225 in C.L. Irwin, P. Garrett, and K. P. McDermott, eds. Proceedings of the 2003 International
Conference on Ecology and Transportation. Center for Transportation and the Environment, North Carolina State
University, Raleigh, N.C. PDF
Lloyd, J. D., and T. E. Martin. 2003. Sibling competition and the evolution of developmental rates. Proceedings of the
Royal Society of London, Series B 270:735-740. (cover article) PDF
Kirkpatrick, C., R.W. Mannan, S. DeStefano, and J. D. Lloyd. 2002. Trends in abundance of grassland birds following a
spring prescribed burn in southern Arizona. Southwestern Naturalist 47:282-292.*
Tewksbury, J. J., and J. D. Lloyd. 2001. Positive interactions under nurse plants: spatial scale, stress gradients, and
benefactor quality. Oecologia 127:425-434. PDF
Lloyd, J.D., R.W. Mannan, S. DeStefano, and C. Kirkpatrick. 1998. The effects of mesquite invasion on a southeastern
Arizona grassland bird community. Wilson Bulletin 110:403-408. PDF
* This paper received the 2002 George Miksch Sutton Award for Conservation Research from the Southwestern
Association of Naturalists.
Some of Dr. Lloyd’s publications are available below in PDF format. For a complete list of publications and
presentations, see Dr. Lloyd’s CV.
Lloyd, J. D., and G. L. Slater. 2007. Environmental factors affecting productivity of brown-headed nuthatches.
Journal of Wildlife Management 71:1968-1975. PDF
Lloyd, J. D., and J. J. Tewksbury. 2007. Analyzing avian nest survival in forests and grasslands: a comparison of the
Mayfield and logistic-exposure methods. Studies in Avian Biology 34:96-104. PDF
Tewksbury, J. J., L. Garner, S. Garner, J. D. Lloyd, V. Saab, and T. E. Martin. 2006. Tests of landscape influence: nest
predation and brood parasitism in fragmented ecosystems. Ecology 87:759-76. PDF
Lloyd, J. D., and A. Casey. 2005. Wildlife hot spots along highways in northwestern Oregon. Final Report to the
Oregon Department of Transportation. PDF
Lloyd, J. D. and T.E. Martin. 2005. Reproductive success of Chestnut-collared Longspurs in native and exotic
grassland. Condor 107:363-374. PDF
Lloyd, J. D., and T. E. Martin. 2004. Nest-site preference and maternal effects on offspring growth. Behavioral
Ecology 15:816-823. PDF
Ruediger, B., and J. D. Lloyd. 2003. A rapid assessment protocol for identifying potential habitat linkages along
roads. Pages 205-225 in C.L. Irwin, P. Garrett, and K. P. McDermott, eds. Proceedings of the 2003 International
Conference on Ecology and Transportation. Center for Transportation and the Environment, North Carolina State
University, Raleigh, N.C. PDF
Lloyd, J. D., and T. E. Martin. 2003. Sibling competition and the evolution of developmental rates. Proceedings of the
Royal Society of London, Series B 270:735-740. (cover article) PDF
Kirkpatrick, C., R.W. Mannan, S. DeStefano, and J. D. Lloyd. 2002. Trends in abundance of grassland birds following a
spring prescribed burn in southern Arizona. Southwestern Naturalist 47:282-292.*
Tewksbury, J. J., and J. D. Lloyd. 2001. Positive interactions under nurse plants: spatial scale, stress gradients, and
benefactor quality. Oecologia 127:425-434. PDF
Lloyd, J.D., R.W. Mannan, S. DeStefano, and C. Kirkpatrick. 1998. The effects of mesquite invasion on a southeastern
Arizona grassland bird community. Wilson Bulletin 110:403-408. PDF
* This paper received the 2002 George Miksch Sutton Award for Conservation Research from the Southwestern
Association of Naturalists.
John is the senior research ecologist at Ecostudies Institute, and manages our Vermont field office.
Contact Dr. John Lloyd
Contact Dr. John Lloyd